NERC funded PhD position on the genetic basis of adaptation to altitude in Heliconius butterflies

27 November 2024

About the Project

We are seeking an enthusiastic student with interests in genetics, ecology and evolution to work on a project investigating thermal adaptation in tropical Andean butterflies. Understanding organisms’ adaptation to their thermal environment is important for predicting responses to climate change. Tropical insects make up around half of all species on Earth, and yet very little is known about their thermal ecology. Butterflies are one of the best-studied insect groups with respect to thermal adaptation, but relatively little is known about the responses of tropical species to climate change.

The Heliconius butterflies have been extensively studied, with good genomic resources and excellent information on species ranges. Many of the species in the genus are found in and around the Andes and work to date shows variation, both within and between species, in traits related to altitudinal and thermal adaptation. This provides an excellent opportunity to investigate the genetic basis of this variation and to find out how genotypes associated with thermal adaptation vary in space and time.

The project can be tailored to the interests of the student but would include analysing genomic data to identify genetic loci associated with altitudinal and thermal adaptation in Heliconius. This could be followed with examination of the distribution of particular genotypes at these loci across South America in relation to different ecological variables and how these have changed over time. This would make use of the extensive existing samples available for Heliconius in various collections, and the genomic data available for many of these. There would also be the possibility of field work in South America for further sampling or experiments. The student would therefore receive training and gain skills in advanced genomics and bioinformatics as well as a range of other ecological and evolutionary techniques.

Co-supervisors: Prof Jon Slate (School of Biosciences), Prof Jane Hill (University of York, Department of Biology), Dr Caroline Bacquet (Amazonian Regional University, IKIAM, Ecuador)

About the funding

This studentship is offered as part of the NERC ACCE+ DLA. ACCE+ is a partnership between the Universities of Sheffield, Liverpool and York, the UK Centre for Ecology and Hydrology (CEH), and the Natural History Museum (NHM), providing doctoral training in the biological components of the natural environment and related disciplines. The ACCE+ DLA is committed to recruiting extraordinary future scientists regardless of age, disability, ethnicity, gender, gender identity, sexual orientation, faith or religious belief, pregnancy or maternity, parental or caring responsibilities or career pathway to date. We understand that a student’s potential can be shown in many ways and we strive to recruit students from all backgrounds, and support them on their scientific journey.

We have designed our application systems to identify candidates who are likely to be successful in research regardless of what opportunities may have been available to them prior to their application.

What is on offer

NERC ACCE+ DLA programme starts from October 2025.

UKRI provide the following funding for 3.5 years:

• Stipend (2024/25 UKRI rate £19,237)

• Tuition Fees at UK fee rate (2024/25 rate £4,786)

• Research support and training grant (RTSG)

Note - UKRI funding only covers UK (Home) fees. The DLA partners have various schemes which allow international students to join the DLA but only be required to pay home fees. Home fees are already covered in the UKRI funding, meaning that successful international candidates do not need to find any additional funding for fees.

International candidates should ensure that they have sufficient funds to pay for a UK student visa, NHS health surcharge fees and travel expenses to the UK, if they are successfully offered a studentship following a formal interview. These costs are not covered by ACCE+.

Not all projects will be funded; the DTP will appoint a limited number of candidates via a competitive process.


All ACCE+ PhDs are available as part time or full time, with part time being a minimum of 50% of full time. Please discuss potential part time arrangements with the primary supervisor before applying to the programme. 

How to apply

Please see the ACCE website for all details of how to apply to the programme at each ACCE+ institution: https://accedtp.ac.uk/how-to-apply/.

Your application should include:

The proforma personal statement, CV and referee details should be submitted through the University of Sheffield Postgraduate Online Application Form. 

Closing date for all applications is Wednesday 8th January 2025 

Candidate webinar

The project primary supervisor will hold a candidate google meet webinar on Tuesday 17th December 2024 at 12-1pm (UK time) to discuss the project with interested candidates. To register for this webinar please complete the google form here